US journalist Evan Gershkovich, of the Wall Street Journal, has been formally arrested in Russia and accused of spying.

Mr Gershkovich, an experienced Russia reporter, was working in Yekaterinburg at the time of his detention.

The Wall Street Journal said it was "deeply concerned" for his safety and vehemently denied the allegations against him.

The Kremlin claimed the reporter had been "caught red-handed".

The FSB said it had "halted illegal activities" and that the reporter had been "acting on US instructions" and "collecting state secrets".

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